Bachelor of Science, The College of William and Mary, Williamsburg VA, 1991-1995
PhD in Molecular Biology, Princeton University, 1995-2001
Thesis: Catherine D. Unrine, 2001: "The Dosage Requirement for Pax6 in Mammalian Lens Development"
Supervisor: Shirley M. Tilghman
Post-doctoral Fellow, Stanford University, Department of Genetics, 2001-2006
Supervisor: Gregory S. Barsh
Assistant Professor, Medical Genetics, University of British Columbia, 2006-2013
Associate Professor, Medical Genetics, University of British Columbia, 2013-present
Canadian Institutes of Health Research New Investigator, 2006-2011
Michael Smith Foundation for Health Research Scholar, 2011-2019